2001 in paleontology

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Paleontology, palaeontology or palæontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.^[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised faeces (coprolites), palynomorphs and chemical residues. Because mankind has encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred in the year 2001.

Arthropods

Newly named insects

Name	Status	Authors	Age	Unit	Location	Notes	Images
Electromyrmococcus^[2]	Valid	Williams	Burdigalian (Miocene)	Dominican amber	Dominican Republic
Tainosia^[3]	Valid	Szwedo Stroiński	Burdigalian (Miocene)	Dominican amber	Dominican Republic

Fishes

Newly named bony fishes

Name	Status	Authors	Age	Unit	Location	Notes	Images
Achoania^[4]	Valid	Zhu Yu Ahlberg	Early Devonian	Xitun Formation	China	The type species is Achoania jarvikii.
Westollrhynchus^[5]	Valid	Schultze	Early Devonian	Bear Rock Formation	Canada	A new genus for "Speonesydrion" lehmanni.

Amphibians

Newly named amphibians

Name	Status	Authors	Age	Unit	Location	Notes
Mesophryne^[6]	Valid	Gao Shubin	Late Jurassic	Yixian Formation	China	The type species is Mesophryne beipiaoensis.
Nezpercius^[7]	Valid	Blob Carrano Rogers Forster Espinoza	Upper Cretaceous	Judith River Formation	USA	The type species is Nezpercius dodsoni.
Sinerpeton^[6]	Valid	Gao Shubin	Late Jurassic	Zhangjiakou Formation	China	The type species is Sinerpeton fengshanensis.

Anapsids

Newly named turtles

Name	Status	Authors	Age	Unit	Location	Notes
Caribemys^[8]	Valid	de la Fuente Iturralde-Vinent	Upper Jurassic	Jagua Formation	Cuba	The type species is Caribemys oxfordiensis.
Cearachelys^[9]	Valid	Gaffney Campos Hirayama	Early Cretaceous	Santana Formation	Brazil	The type species is Cearachelys placidoi.
Kurmademys^[10]	Valid	Gaffney Chatterjee Rudra	Late Cretaceous	Kallamedu Formation	India	The type species is Kurmademys kallamedensis.

Ichthyosaurs

Name	Status	Authors	Age	Unit	Location	Notes	Images
Metashastasaurus^[11]	Synonymy	Nichollis Manabe	Late Triassic	Pardonet Formation	Canada	A new genus for "Shastasaurus" neoscapularis. A junior synonymy of Callawayia Maisch & Matzke, 2000.

Lepidosauromorphs

Newly named basal lepidosauromorphs

Name	Status	Authors	Age	Unit	Location	Notes	Images
Hypuronector^[12]	Valid	Colbert Olsen	Late Triassic	Lockatong Formation	USA	The type species is Hypuronector limnaios.

Newly named plesiosaurs

Name	Status	Authors	Age	Unit	Location	Notes	Images
Hauffiosaurus^[13]	Valid	O'Keefe	Lower Jurassic	Posidonia Shale Formation	Germany	The type species is Hauffiosaurus zanoni.

Newly named sphenodonts

Name	Status	Authors	Age	Unit	Location	Notes	Images
Godavarisaurus^[14]	Valid	Evans Prasad Manhas	Early Jurassic	Kota Formation	India	The type species is Godavarisaurus latefi.
Rebbanasaurus^[14]	Valid	Evans Prasad Manhas	Early Jurassic	Kota Formation	India	The type species is Rebbanasaurus jani.

Archosauromorphs

Newly named basal archosauromorphs

Name	Status	Authors	Age	Unit	Location	Notes	Images
Yonghesuchus^[15]	Valid	Wu Liu Li	Upper Triassic	Tongchuan Formation	China	The type species is Yonghesuchus sangbiensis.

Newly named crurotarsans

Name	Status	Authors	Age	Unit	Location	Notes
Pabwehshi^[16]	Valid	Wilson Malkani Gingerich	Upper Cretaceous	Pab Formation	Pakistan	The type species is Pabwehshi pakistanensis.
Rugosuchus^[17]	Valid	Wu Wu Russell	Lower Cretaceous	?Nenjiang Formation	China	The type species is Rugosuchus nonganensis.
Stratiotosuchus^[18]	Valid	Campos Suarez Riff Kellner	Upper Cretaceous	Adamantina Formation	Brazil	The type species is Stratiotosuchus maxhechti.

Newly named dinosaurs

Data courtesy of George Olshevsky's dinosaur genera list^[19] and Dr. Jeremy Montague's dinosaur genus database.^[20]

Name	Status	Authors	Unit	Location	Notes
"Alashansaurus"^[21]	Nomen ex dissertatione	Daniel Chure	Ulansuhai Formation	China	Would later be formally named Shaochilong in 2009.
Aletopelta^[22]	Valid taxon	Ford James Kirkland	Point Loma Formation	USA	A 20-foot-long (6.1 m) ankylosaurid. Apparently, before being fossilized, the animal's bloated carcass had floated out to sea and formed a miniature reef environment after it sunk to the bottom.
Bienosaurus^[23]	Valid taxon	Dong Zhiming	Lower Lufeng Formation	China	A primitive scelidosaurid known from scant remains.
Cedarpelta^[24]	Valid taxon	Kenneth Carpenter James Kirkland et al.^{[CAL 1]}	Cedar Mountain Formation	USA	The most basal known ankylosaurid.
Citipati^[25]	Valid taxon	James M. Clark Mark Norell Rinchen Barsbold	Djadokhta Formation	Mongolia	A relatively large oviraptorid known to brood its nests.
Draconyx^[26]	Valid taxon	Octávio Mateus Antunes	Lourinhã Formation	Portugal	A relative of Camptosaurus.
Eotyrannus^[27]	Valid taxon	Hutt Naish et al.^{[CAL 2]}	Wessex Formation	UK	A twenty foot tyrannosauroid.
Eshanosaurus^[28]	Valid taxon	Xu Xing Zhao Xijin James M. Clark	Lower Lufeng Formation	China	A therizinosauroid and possibly the earliest known coelurosaur.
Gobisaurus^[29]	Valid taxon	Matthew K. Vickaryous Anthony P. Russell et al.^{[CAL 3]}	Ulansuhai Formation	China	An ankylosaurid that resembled Shamosaurus.
“Hanwulosaurus”^[30]	Nomen nudum	Anonymous		China	A thirty foot ankylosaur.
"Heilongjiangosaurus"^[31]	Nomen nudum	Li W. Jin J.		China	A hadrosaur, possibly synonymous with Charonosaurus.
Hesperosaurus^[32]	Valid taxon	Kenneth Carpenter Miles Cloward	Morrison Formation	USA	A stegosaurid slightly older and more primitive than Stegosaurus, although the genera may be synonymous.
Jiangshanosaurus^[33]	Valid taxon	Tang F. Kang et al.^{[CAL 4]}		China	A titanosaur known only from a partial skeleton found near Lixian Village, China.
Jinzhousaurus^[34]	Valid taxon	Wang X. Xu Xing	Yixian Formation	China	A hadrosauroid known from a nearly complete skeleton.
Khaan^[25]	Valid taxon	James M. Clark Mark Norell Rinchen Barsbold	Djadochta Formation	Mongolia	A fairly typical oviraptorid once misidentified as Ingenia.
"Kittysaurus"	Junior synonym of Eotyrannus	Hargreaves	Wessex Formation	UK
Liaoningosaurus^[35]	Valid taxon	Xu Xing Wang X. You	Yixian Formation	China	A bizarre ankylosaur of uncertain classification. Known from the complete specimen of a juvenile 34 cm long, it's the smallest known ankylosaur to date.
Losillasaurus^[36]	Valid taxon	Maria Lourdes Casanovas-Cladellas Santafe Sanz		Spain	A large turiasaur known from a partial subadult skeleton.
Masiakasaurus^[37]	Valid taxon	Sampson Carrano Forster	Maevarano Formation	Madagascar	A 2 meter (about 6–7 feet) noasaurid with unusual forward-pointing teeth.
Megapnosaurus^[38]	Same as Syntarsus.	Ivie Slipinski Wegrzynowicz			Formerly known as Syntarsus, this new name was erected by entomologists due to synonymy between the dinosaur and a genus of beetle.
Neimongosaurus^[39]	Valid taxon	Zhang X. H. Xu Xing et al.^{[CAL 5]}	Iren Dabasu Formation	China	A therizinosaur about 2.3 meters in length.
Nothronychus^[40]	Valid taxon	James Kirkland Wolfe	Moreno Hill Formation	USA	A therizinosaur.
Paralititan^[41]	Valid taxon	J. B. Smith Lamanna et al.^{[CAL 6]}	Bahariya Formation	Egypt	A titanosaur and of the most massive dinosaurs ever discovered, with an estimated weight of 59 tonnes (65 short tons) and length of around 26 meters (85 ft).
Planicoxa^[42]	Valid taxon	DiCroce Kenneth Carpenter	Cedar Mountain Formation	USA	An advanced iguanodontian.
Pukyongosaurus^[43]	Valid taxon	Dong Zhiming Paik Kim H. J.	Hasandong Formation	South Korea	A titanosauriform related to Euhelopus.
Quilmesaurus^[44]	Valid taxon	Rodolfo Coria	Allen Formation	Argentina	A 5–6 meter (16–20 feet) theropod known from a partial leg.
Rapetosaurus^[45]	Valid taxon	Kristina Curry-Rogers Catherine A. Forster	Maevarano Formation	Madagascar	A 15 metres (49 ft) titanosaur.
Ruehleia^[46]	Valid taxon	Peter Galton.	Knollenmergel	Germany	A prosauropod named for Hugo Ruehle von Lilienstern.
"Szechuanoraptor"^[21]	Nomen ex dissertatione	Daniel Chure	Dashanpu Formation	China	A theropod not yet formally described. The name was coined by Daniel Chure in 2000.
Wellnhoferia^[47]	Valid non-dinosaurian taxon.	Andrzej Elżanowski	Solnhofen Limestone	Germany	Very similar to Archaeopteryx, which may be its senior synonym.
Venenosaurus^[48]	Valid taxon	Tidwell Kenneth Carpenter S. Meyer	Cedar Mountain Formation	USA	A relatively small (probably around 10 m (33 ft) long) titanosauriform sauropod, known from an incomplete skeleton of an adult and a juvenile. Its tail vertebrae articulated in a unique fashion that may be of evolutionary significance.
"Yibinosaurus"^[49]	Nomen nudum	Ouyang vide: Anonymous		China	A saropod from Szechuan slated to be described by Ouyang Hui.

Laelaps trihedrodon synonymized with Allosaurus

Laelaps trihedrodon was one of the many species described by Edward Drinker Cope that he referred to his preoccupied genus Laelaps, later renamed Dryptosaurus. Although the type specimen included a partial dentary, all material except for a collection of five damaged partial tooth crowns (AMNH 5780) has been lost.^[50] The rediscovered material had lately been mistaken by Mcintosh for the type specimen in 1998. However this identification was impossible because the type's teeth were described by Cope as "smooth" and having a "fine silky luster," while the AMNH 5780 teeth were thoroughly worn and had matrix clinging to them. Most of the type specimen's teeth were successional, but all the AMNH 5780 teeth were functional.^[51] AMNH 5780 has many features in common with Allosaurus and is probably referrable to that genus. However some of the Allosaurus-like characters of the tooth are primitive to theropods as a whole and may have been present in the less studied or poorly preserved Morrison theropod species. Consequently the synonymization of L. trihedrodon with Allosaurus is tentative, despite its high likelihood.^[52]

Carcharodontosaurus saharicus brain and inner ear examined

Hans C. E. Larsson published a description of the inner ear and endocranium of Carcharodontosaurus saharicus.^[53] The C. saharicus braincase "completely encloses the endocranial region." This "high degree of ossification" made detailed analysis of its anatomy significantly easier.^[54] The C. saharicus endocast is similar to that of a related dinosaur, Allosaurus fragilis. Larsson describes the olfactory bulbs and peduncles as lying "on approximately the same horizontal plane as the forebrain." The cephalic flexure, the bend between the fore- and midbrain, has an angle of 45 degrees. The pontine flexure, the bend between the mid- and hindbrain has an angle of about 40 degrees. Carcharodontosaurus had a large optic (II) nerve. The C. saharicus vena capitis dorsalis" drains the anterior neck muscles through a pair of long canals on the posterior surface of the endocast." Allosaurus, Dromaeosaurus albertensis, although in C. saharicus and Troodon "the transverse sinus probably drained into a middle cerebral vein that exited the brain in the ridge present on the dorsal edge of the trigeminal foramen."

The three semicircular canals of the inner ear of Carcharodontosaurus saharicus, when viewed from the side, had a subtriangular outline. This subtriangular inner ear configuration is present in Allosaurus, lizards, turtles, but not in birds. The pointed apex "at the junction of the anterior and posterior semicircular canals" is caused by the near linearity of the canals and closely resembles the condition of modern crocodiles. The subtriangular configuration may be the basal condition of archosauromorphs. A recess which would have held the floccular lobe of the brain projects into the area surrounded by the semicircular canals. This condition is also present in other non-avian theropods, birds, and pterosaurs. The orientation of the lagena of C. saharicus resembles the condition in crocodillians and some birds. The extent of its perilymphatic duct resembled those of Varanus, crocodillians, and birds. The crista which would have supported the secondary tympanic membrane in C. saharicus was either absent, or not preserved. This contrasts with Troodon, whose crista were ossified at least in their dorsal and ventral regions and their remaining portions either cartilaginous or too delicate to be preserved. The metotic strut of C. saharicus is reduced and medial compared to the "laterally hypertrophied" condition of non-avian maniraptors like Dromaeosaurus and Troodon, as well as primitive birds like Archaeopteryx and Hesperornis.

Newly named birds

Name	Status	Authors	Age	Unit	Location	Notes
Apsaravis^[55]	Valid	Norell Clarke	Late Cretaceous	Djadokhta Formation	Mongolia	The type species is Apsaravis ukhaana.
Limenavis^[56]	Valid	Clarke Chiappe	Late Cretaceous	Allen Formation	Argentina	The type species is Limenavis patagonica.
Longipteryx^[57]	Valid	Zhang Zhou Hou Gu	Early Cretaceous	Jiufotang Formation	China	The type species is Longipteryx chaoyangensis.
Yanornis^[58]	Valid	Zhou Zhang	Early Createcous	Jiufotang Formation	China	The type species is Yanornis martini.
Yixianornis^[58]	Valid	Zhou Zhang	Early Cretaceous	Jiufotang Formation	China	The type species is Yixianornis grabaui.

Newly named pterosaurs

Name	Status	Authors	Age	Unit	Location	Notes	Images
Haopterus^[59]	Valid	Wang Lü	Early Cretaceou	Yixian Formation	China	The type species is Haopterus gracilis.
Istiodactylus^[60]	Valid	Howse Milner Martill	Early Cretaceous	Vectis Formation	United Kingdom	A new genus for Ornithodesmus latidens.

Synapsids

Non-mammalian

Name	Status	Authors	Age	Unit	Location	Notes
Delectosaurus^[61]	Valid	Kurkin	Late Permian	Vyazniki Assemblage	Russia	A dicynodont. Two species are described Delectosaurus arefjevi and D. berezhanensis.
Interpresosaurus^[61]	Valid	Kurkin	Late Permian	Vyazniki Assemblage	Russia	A dicynodont. The type species is Interpresosaurus blomi.
Lumkuia^[62]	Valid	Hopson Kitching	Middle Triassic	Burgersdorp Formation	South Africa	A cynodont. The type species is Lumkuia fuzzi.
Malasaurus^[63]	Valid	Tatarinov	Late Triassic	Vyazniki Assemblage	Russia	A therocephalian. The type species is Malasaurus germanus.
Mitredon^[64]	Valid	Shapiro Jenkins	Late Triassic	Fleming Fjord Formation	Greenland	A cynodont. The type species is Mitredon cromptoni.
Prozostrodon^[65]	Valid	Bonaparte Barberena	Late Triassic	Santa Maria Formation	Brazil	New genus for "Thrinaxodon" brasiliensis Barberena, Bonaparte & Sá Teixeira, 1987
Riograndia^[66]	Valid	Bonaparte Ferigolo Ribeiro	Late Triassic	Caturrita Formation	Brazil	A cynodont. The type species is Riograndia guaibensis.

Plants

Pteridophyta

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Wessiea^[67]^[68]	gen et sp nov	Valid	Pigg & Rothwell	Langhian (Middle Miocene)	"Ho ho" site, Grande Ronde Basalt, Columbia River Basalt Group.	USA

Conifers

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Metasequoia foxii^[69]	sp nov	Valid	Stockey, Rothwell, & Falder	late Tiffanian	Munce’s Hill and Gao mine sites, Paskapoo Formation	Canada

Angiosperms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Trochodendron nastae^[70]	sp nov	Valid	Pigg, Wehr, & Ickert-Bond	Ypresian	Tom Thumb Tuff, Klondike Mountain Formation	USA

Footnotes

Complete author list

As science becomes more collaborative, papers with large numbers of authors are becoming more common. To prevent the deformation of the tables, these footnotes list the contributors to papers that erect new genera and have many authors.

^ Kenneth Carpenter, James Kirkland, Burge, Bird.
^ Hutt, Naish, Martill, Barker, Newberry.
^ Matthew K. Vickaryous, Anthony P. Russell, Phillip Currie, Zhao Xijin.
^ Tang F., Kang, Jin X., Wei, Wu W. T.
^ Zhang X. H., Xu Xing, Zhao X., Paul Sereno, Kuang, Tan L.
^ J. B. Smith, Lamanna, Lacovara, Dodson, J. R. Smith, Poole, Giegengack, Attia.

References

^ Newman, Garfield, et al (2001). Echoes from the past: world history to the 16th century. Toronto: McGraw-Hill Ryerson Ltd. ISBN 0-07-088739-X.
^ Johnson, M.S.; et al (2001). "Acropyga and Azteca Ants (Hymenoptera: Formicidae) with Scale Insects (Sternorrhyncha: Coccoidea): 20 Million Years of Intimate Symbiosis". American Museum Novitates 3335: 1–18. doi:10.1206/0003-0082(2001)335<0001:AAAAHF>2.0.CO;2.
^ Szwedo, J.; Stroiński, A. (2001). "Tainosia quisqueyae gen. and sp. nov. from the Oligocene/Miocene Dominican amber (Hemiptera: Fulgoroidea: Nogodinidae)". Genus 12 (1): 29–34. http://ag.udel.edu/delpha/5325.pdf.
^ Zhu, M, Yu, X. & Ahlberg, P.E. (2001). "A primitive sacropterygian fish with an eyestock". Nature 410: 81–84. doi:10.1038/35065078.
^ Schultze, H.-P. (2001). "Melanognathus, a primitive dipnoan from the Lower Devonian of the Canadian Arctic and the interrelationships of Devonian dipnoans". Journal of Vertebrate Paleontology 21 (4): 781–794.
^ ^a ^b Gao, K.-Q. & Shubin, N.H. (2001). "Late Jurassic salamanders from northern China". Nature 410 (6828): 574–577. Bibcode 2001Natur.410..574G. doi:10.1038/35069051. PMID 11279493.
^ Blob, R.W., Carrano, M.T., Rogers, R.R., Forster, C.A. & Espinoza, N.R. (2001). "A new fossil frog from the Upper Cretaceous Judith River Formation of Montana". Journal of Vertebrate Paleontology 21 (1): 190–194. doi:10.1671/0272-4634(2001)021[0190:ANFFFT]2.0.CO;2.
^ de la Fuente, M.S. & Iturralde-Vinent, M. (2001). "A new pleurodiran turtle from the Jagua Formation (Oxfordian) of western Cuba". Journal of Paleontology 75 (4): 860–869.
^ Gaffney, E.S., de Almedia Campos, D. & Hirayama, R. (2001). "Cerachelys, a new side-necked turtle (Pelomedusoides: Bothremydidae) from the Early Cretaceous of Brazil". American Museum Novitates (3319): 1–20.
^ Gaffney, E.S., Chatterjee, S., Rudra, D.K. (2001). "Kurmademys, a new side-necked turtle (Pelomedusoides: Bothremydidae) from the Late Cretaceous of India". American Museum Novitates (3321).
^ Nichollis, E.L. & Manabe, M. (2001). "A new genus of ichthyosaur from the Late Triassic Pardonet Formation of British Columbia: bridging the Triassic-Jurassic gap". Canadian Journal of Earth Science 38: 983–1002.
^ Colbert, E.H. & Olsen, P.E. (2001). "A new and unusual aquatic reptile from the Lockatong Formation of New Jersey (Late Triassic, Newark Supergroup)". American Museum Novitates 3334: 1–15.
^ O'Keefe, F.R. (2001). "A cladistic analysis and taxonomic revision of the Plesiosauria (Reptilia: Sauropterygia)". Acta Zoologica Fennica 213: 1–63.
^ ^a ^b Evans, S.E., Prasad, G.V.R. & Manhas, B.K. (2001). "Rhynchocephalians (Diapsida: Lepidosauria) from the Jurassic Kota Formation of India". Zoological Journal of the Linnean Society 133: 309–334.
^ Wu, X.-C., Liu, J. & Li, J.-L. (2001). "The anatomy of the first archosauriform (Diapsida) from the terrestrial Upper Triassic of China". Vertebrata PalAsiatica 39 (4): 251–265.
^ Wilson, J. A., Malkani, M.S. & Gingerich, P.D. (2001). "New crocodyliform (Reptilia, Mesoeurocodylia) from the Upper Cretaceous Pab Formation of Vitakri, Balochistan (Pakistan)". Contributions from the Museum of Paleontology, The University of Michigan 30 (12): 321–336.
^ Wu, X.-C., Wu, Z.-W. & Russell, A.P. (2001). "Cranial anatomy of a new crocodyliform (Archosauria: Crocodylomorpha) from the Lower Cretaceous of Song-Liao Plain, northeastern China". Canadian Journal of Earth Science 38 (12): 1653–1663.
^ Campos, D. de A., Suarez, J.M., Riff, D. & Kellner, A.W.A. (2001). "Short note on a new Baurusuchidae (Crocodyliformes, Metasuchia) from the Upper Cretaceous of Brazil". Boletim do Museu Nacional, Geologia (57): 1–7.
^ Olshevsky, George. "Dinogeorge's Dinosaur Genera List". http://www.polychora.com/dinolist.html. Retrieved 2008-08-07.
^ Montague, Jeremy. "Dr. Montague's Database". http://www.barry.edu/bio300/pdf/montague%20dino%20web%20data%2012-3-2005.pdf. Retrieved 2008-10-29.
^ ^a ^b Chure, D. 2000. A new species of Allosaurus from the Morrison Formation of Dinosaur National Monument (UT-CO) and a revision of the theropod Family Allosauridae. Note: Unpublished Ph.D. dissertation.
^ Ford, T.L. and J.I. Kirkland. 2001. Carlsbad ankylosaur (Ornithischia, Ankylosauria): an ankylosaurid and not a nodosaurid. Chapter 12 (pp. 239-260) in: The Armored Dinosaurs (K. Carpenter, ed.). Indiana University Press, Bloomington & Indianapolis, IA (483 pages): 239-260.
^ Dong Z. 2001. Primitive Armored Dinosaur from the Lufeng Basin, China. pp. 237-242 In: Mesozoic Vertebrate Life: New Research inspired by the Paleontology of Philip J. Currie (D.H. Tanke and K. Carpenter, eds.). Indiana University Press, Bloomington & Indianapolis, IA: xviii + 542 pages.
^ Carpenter, K., J.I. Kirkland, D. Burge, and J. Bird. 2001. Disarticulated Skull of a New Primitive Ankylosaurid from the Lower Cretaceous of Eastern Utah. Chapter 11 of The Armored Dinosaurs (K. Carpenter, ed.). Indiana University Press, Bloomington & Indianapolis, IA: pp. 211-238.
^ ^a ^b Clark, J.M., M.A. Norell, and P. Barsbold. 2001. Two new oviraptorids (Theropoda: Oviraptorosauria), Upper Cretaceous Djadokhta Formation, Ukhaa Tolgod, Mongolia. Journal of Vertebrate Paleontology 21 (2): pp. 209–213.
^ Mateus, O. and M.T. Antunes. 2001. Draconyx loureiroi, a new Camptosauridae (Dinosauria, Ornithopoda) from the Late Jurassic of Lourinha, Portugal. Annales Paleontol. 87 (1): pp. 61-73.
^ Hutt, S., D. Naish, D.M. Martill, M.J. Barker, and P. Newbery. 2001. A preliminary account of a new tyrannosauroid theropod from the Wessex Formation (Early Cretaceous) of southern England. Cretaceous Research 22 (2): pp. 227-242.
^ Xu, X., X. Zhao, J.M. Clark. 2001. A new therizinosaur from the Lower Jurassic lower Lufeng Formation of Yunnan, China. Journal of Vertebrate Paleontology 21 (3): pp. 477–483.
^ Vickaryous, M.K., A.P. Russell, P.J. Currie, and X.- J. Zhao. 2001. A new ankylosaurid (Dinosauria: Ankylosauria) from the Lower Cretaceous of China, with comments on ankylosaurian relationships. In: The Sino-Canadian Dinosaur Project 3. Canadian Journal of Earth Sciences 38 (12): pp. 1767-1780.
^ Xinhua News Service news articles
^ Li, W. and J. Jin. 2001. On the Upper Cretaceous Jiayin Group of Heilongjiang Province, China (Deng and Wang, eds): 65-74.
^ Carpenter, K., C.A. Miles, and K. Cloward. 2001. New primitive stegosaur from the Morrison Formation, Wyoming. In: The Armored Dinosaurs (Carpenter, K., ed.). Indiana University Press, Bloomington, IA: pp. 55-75.
^ Tang F., X.-M. Kang, X.-S. Jin., F. Wei, and W.- T. Wu. 2001. A new sauropod dinosaur of Cretaceous from Jiangshan, Zhejiang Province. Vertebrate Palasiatica 39 (4): pp. 272-281.
^ Wang, X. and X. Xu. 2001. A new iguanodontid (Jinzhousaurus yangi gen. et sp. nov.) from the Yixian Formation of western Liaoning, China. Chinese Science Bulletin 46 (19): pp. 1669-1672.
^ Xu, X., X.-L. Wang, and H.-L. You. 2001. A juvenile ankylosaur from China [Liaoningosaurus paradoxus]. Die Naturwissenshaften 88 (7): pp 297-300.
^ Casanovas, M.L., J.V. Santafe, and J.L. Sanz. 2001. Losillasaurus giganteus, un nuevo sauropodo del transito Jurasico-Cretacico de la cuenca de "Los Serranos" (Valencia, España). Paleontologia i Evolucio, 32-33: pp. 99-122.
^ Sampson, S. D. M.T. Carrano, and C.A. Forster. 2001. A bizarre predatory dinosaur from the Late Cretaceous of Madagascar. Nature 409: pp. 504-509.
^ Ivie, M.A., S.A. Slipinski, and P. Wegrzynowicz. 2001. Generic homonyms in the Colydiinae (Coleoptera: Zopheridae). Insecta Mundi 15: pp.63-64.
^ Zhang, X-H., X. Xu, P. Sereno, X.-W. Kuang, and L. Tan. 2001. A long-necked therizionosauroid dinosaur from the Upper Cretaceous Iran Dabasu Formation of Nei Mongol, People's Republic of China. Vertebrate Palasiatica 39 (4): pp. 282-290.
^ Kirkland, J.I. and D.G. Wolf. 2001. First definitive therizinosaurid (Dinosauria; theropoda) from North America. Journal of Vertebrate Paleontology 21 (3): pp. 410-414.
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